GEVIBench › Brightness network

Brightness network

Molecular brightness of each sensor relative to EGFP. Sensors measured against another sensor rather than EGFP are placed through the chain of pairwise comparisons below (multiple paths combined by geometric mean).

Brightness relative to EGFP

Pairwise brightness measurements

• 2Photron is 4.12× as bright as JEDI-2P — Villette 2024, Fig. 3f
• Ace2N-mNeon is 2.76× as bright as EGFP — Shaner 2013, Table 1
• Ace2N-mNeon is 0.9090909090909091× as bright as Ace2N-mNeon2 — Kannan 2022, Fig. S11A
• Ace2N-mNeon is 5.988023952095808× as bright as ASAP3 — Villette 2019, Table 1
• Ace2N-mNeon is 5.555555555555555× as bright as FlicR1 — Kannan 2018, Fig. 1j
• Ace2N-mNeon is 5.555555555555555× as bright as FlicR2 — Kannan 2018, Fig. 1j
• Ace2N-mNeon is 1.4285714285714286× as bright as pAce — Kannan 2022, Fig. S11A
• Ace2N-mNeon is 1.4084507042253522× as bright as VARNAM — Kannan 2018, Fig. 1j
• Ace2N-mNeon is 0.25× as bright as Voltron — Abdelfattah 2019, Fig. 1F
• Ace2N-mNeon2 is 1.1× as bright as Ace2N-mNeon — Kannan 2022, Fig. S11A
• Ace2N-mNeon2 is 0.5× as bright as Vega — Cao 2025, Fig. 1B
• Ace2N-mNeon2 is 0.625× as bright as ElectraOFF — Zhang 2025, Fig. 1e
• Arch is 0.00016× as bright as EGFP — Herasymenko 2025, Table 1
• Arch is 0.2× as bright as Archer1 — Flytzanis 2014, Fig. 1a
• Arch is 0.3333333333333333× as bright as Archer2 — Flytzanis 2014, Fig. 1a
• Arch is 0.0006858710562414266× as bright as FlicR1 — Monakhov 2020, Table 1
• Arch is 0.00205761316872428× as bright as nirButterfly — Monakhov 2020, Table 1
• Arch is 0.06578947368421052× as bright as QuasAr1 — Hochbaum 2014, Table S2
• Arch is 0.05× as bright as QuasAr1 — Monakhov 2020, Table 1
• Arch is 0.29411764705882354× as bright as QuasAr2 — Hochbaum 2014, Table S2
• Arch is 0.03571428571428571× as bright as SomArchon — Monakhov 2020, Table 1
• Arch is 0.0004374453193350831× as bright as VARNAM — Monakhov 2020, Table 1
• Archer1 is 1× as bright as QuasAr2 — Monakhov 2020, Table 1
• Archer1 is 5× as bright as Arch — Flytzanis 2014, Fig. 1a
• Archer1 is 0.00367× as bright as EGFP — Nikolaev 2025, Table 2
• Archer2 is 3× as bright as Arch — Flytzanis 2014, Fig. 1a
• Archer2 is 0.0163× as bright as EGFP — Nikolaev 2025, Table 2
• Archon1 is 2.78× as bright as QuasAr2 — Piatkevich 2018, Fig. 2b
• Archon1 is 0.588× as bright as QuasAr6a — Tian 2023, Ext. Fig. 4b
• Archon1 is 0.5× as bright as QuasAr6b — Tian 2023, Ext. Fig. 4b
• Archon1 is 0.005076142131979695× as bright as ASAP3 — Villette 2019, Table 1
• Archon1 is 0.14285714285714285× as bright as nirButterfly — Song 2024, Fig. 2b
• Archon1 is 0.3333333333333333× as bright as NovArch — Chien 2021, Fig. S1C
• Archon2 is 8.01× as bright as QuasAr2 — Piatkevich 2018, Fig. 2b
• ArcLight is 0.655× as bright as EGFP — Villette 2019, Table 1
• ArcLight is 1.639344262295082× as bright as ASAP1 — Milosevic 2020, Fig. 2F
• ArcLight is 1.4662756598240467× as bright as ASAP3 — Villette 2019, Table 1
• ArcLight is 3.4482758620689657× as bright as Bongwoori — Lee 2017, Table 3
• ArcLight is 0.3333333333333333× as bright as MacQ-mCitrine — Gong 2014, Main text
• ArcLight is 0.3333333333333333× as bright as MacQ-mOrange2 — Gong 2014, Main text
• ArcLight is 1.4771048744460855× as bright as Marina — Platisa 2017, Main text
• ASAP1 is 1× as bright as ASAP2s — Chamberland 2017, Fig. S2
• ASAP1 is 0.61× as bright as ArcLight — Milosevic 2020, Fig. 2F
• ASAP1 is 0.94× as bright as ASAP3 — Nair 2025, Fig. 2C
• ASAP2s is 1.11× as bright as ASAP3 — Liu 2022, Fig. 2I
• ASAP2s is 1× as bright as ASAP1 — Chamberland 2017, Fig. 1—figure supplement 2A
• ASAP2s is 0.45454545454545453× as bright as JEDI-1P — Lu 2023, Fig. 2h
• ASAP2s is 0.6666666666666666× as bright as JEDI-2P — Liu 2022, Fig. 2I
• ASAP3 is 0.446× as bright as EGFP — Villette 2019, Table 1
• ASAP3 is 0.682× as bright as ArcLight — Villette 2019, Table 1
• ASAP3 is 197× as bright as Archon1 — Villette 2019, Table 1
• ASAP3 is 556× as bright as QuasAr2 — Villette 2019, Table 1
• ASAP3 is 0.167× as bright as Ace2N-mNeon — Villette 2019, Table 1
• ASAP3 is 0.87× as bright as ASAP2s — Lu 2023, Fig. 2h
• ASAP3 is 556× as bright as paQuasAr3 — Villette 2019, Table 1
• ASAP3 is 1.0638297872340425× as bright as ASAP1 — Nair 2025, Fig. 2C
• ASAP3 is 2.2381378692927485× as bright as ASAP4b — Evans 2023, Ext. Fig. 3d
• ASAP3 is 2.7624309392265194× as bright as ASAP4e — Evans 2023, Ext. Fig. 3d
• ASAP3 is 1× as bright as ASAP5 — Hao 2024, Fig. 1G
• ASAP3 is 0.4098360655737705× as bright as JEDI-1P — Lu 2023, Fig. 2h
• ASAP3 is 0.5847953216374269× as bright as JEDI-1P — Nair 2025, Fig. 2C
• ASAP3 is 0.5988023952095809× as bright as JEDI-2P — Liu 2022, Fig. 2I
• ASAP3 is 0.5592841163310962× as bright as rEstus-NI — Nair 2025, Fig. 2C
• ASAP3 is 0.48309178743961356× as bright as rEstus — Rühl 2024, Table 1
• ASAP3 is 0.5847953216374269× as bright as rEstus — Nair 2025, Table S1
• ASAP3 is 0.9276437847866419× as bright as rEstus2s — Rühl 2026, Table S1
• ASAP4b is 0.4468× as bright as ASAP3 — Evans 2023, Ext. Fig. 3d
• ASAP4e is 0.362× as bright as ASAP3 — Evans 2023, Ext. Fig. 3d
• ASAP4e is 0.5× as bright as FORCE1s — McDonald 2026, Fig. 1C
• ASAP4e is 0.6666666666666666× as bright as ASAP5 — Lee 2024, Table S4
• ASAP4e is 1× as bright as ASAP6c — Lee 2024, Table S4
• ASAP4e is 0.03571428571428571× as bright as pAce — Lee 2024, Table S4
• ASAP4e is 0.2857142857142857× as bright as Positron — Lee 2024, Table S4
• ASAP5 is 1× as bright as ASAP3 — Hao 2024, Fig. 1G
• ASAP5 is 1.5× as bright as ASAP4e — Lee 2024, Table S4
• ASAP5 is 0.78125× as bright as ASAP7y — Hao 2026, Fig. 2l
• ASAP5 is 0.6896551724137931× as bright as JEDI-2P — Land 2026, Ext. Fig. 1c
• ASAP5 is 0.8× as bright as JEDI-3hyp — Land 2026, Ext. Fig. 1c
• ASAP5 is 0.7194244604316548× as bright as JEDI-3sub — Land 2026, Ext. Fig. 1c
• ASAP6c is 1× as bright as ASAP4e — Lee 2024, Table S4
• ASAP7y is 1.28× as bright as ASAP5 — Hao 2026, Fig. 2l
• Bongwoori is 0.29× as bright as ArcLight — Lee 2017, Table 3
• Bongwoori is 0.5291005291005292× as bright as Bongwoori-R3 — Lee 2017, Table 3
• Bongwoori-R3 is 1.89× as bright as Bongwoori — Lee 2017, Table 3
• CAESR is 1.428× as bright as EGFP — Zou 2014, Table S1
• Cepheid1b is 1.65× as bright as VARNAM — Han 2023, Fig. S5
• Cepheid1s is 0.43× as bright as VARNAM — Han 2023, Fig. S5
• ElectraOFF is 1.6× as bright as Ace2N-mNeon2 — Zhang 2025, Fig. 1e
• ElectraON is 1.3× as bright as pAce — Zhang 2025, Fig. 1e
• ElectricPk is 0.32× as bright as EGFP — doi:estimated from cpEGFP (GCaMP3) EC×QY vs EGFP
• FlicR1 is 1458× as bright as Arch — Monakhov 2020, Table 1
• FlicR1 is 0.18× as bright as Ace2N-mNeon — Kannan 2018, Fig. 1j
• FlicR2 is 0.18× as bright as Ace2N-mNeon — Kannan 2018, Fig. 1j
• FlicR2 is 1.1764705882352942× as bright as VADER1 — Yang 2026, Fig. 2C
• FORCE1s is 0.39× as bright as JEDI-2P — McDonald 2026, Fig. 1C
• FORCE1s is 2× as bright as ASAP4e — McDonald 2026, Fig. 1C
• HArclight1 is 1.808× as bright as EGFP — Deo 2021, Table S4
• HASAP1 is 1.808× as bright as EGFP — Deo 2021, Table S4
• HVI is 0.19× as bright as Voltron — Liu 2021, Table S4
• HVI is 1.8867924528301885× as bright as HVI+ — Liu 2025, Fig. S5C
• HVI+ is 0.53× as bright as HVI — Liu 2025, Fig. S5C
• Jarvis is 0.42× as bright as pAce — Grimm 2026, Fig. 1C
• Jarvis is 4.08× as bright as JEDI-2P — Grimm 2026, Fig. 1C
• JEDI-1P is 2.44× as bright as ASAP3 — Lu 2023, Fig. 2h
• JEDI-1P is 2.2× as bright as ASAP2s — Lu 2023, Fig. 2h
• JEDI-1P is 1.71× as bright as ASAP3 — Nair 2025, Fig. 2C
• JEDI-1P is 0.125× as bright as Vega — Cao 2025, Fig. 1B
• JEDI-1P is 1.3550135501355014× as bright as rEstus2s — Rühl 2026, Table S1
• JEDI-2P is 1.67× as bright as ASAP3 — Liu 2022, Fig. 2I
• JEDI-2P is 1.5× as bright as ASAP2s — Liu 2022, Fig. 2I
• JEDI-2P is 1.45× as bright as ASAP5 — Land 2026, Ext. Fig. 1c
• JEDI-2P is 0.24271844660194175× as bright as 2Photron — Villette 2024, Fig. 3f
• JEDI-2P is 2.564102564102564× as bright as FORCE1s — McDonald 2026, Fig. 1C
• JEDI-2P is 0.24509803921568626× as bright as Jarvis — Grimm 2026, Fig. 1C
• JEDI-2P is 1.1627906976744187× as bright as JEDI-3hyp — Land 2026, Fig. 1d
• JEDI-2P is 1.0416666666666667× as bright as JEDI-3sub — Land 2026, Fig. 1d
• JEDI-3hyp is 0.86× as bright as JEDI-2P — Land 2026, Fig. 1d
• JEDI-3hyp is 1.25× as bright as ASAP5 — Land 2026, Ext. Fig. 1c
• JEDI-3sub is 0.96× as bright as JEDI-2P — Land 2026, Fig. 1d
• JEDI-3sub is 1.39× as bright as ASAP5 — Land 2026, Ext. Fig. 1c
• MacQ-mCitrine is 3× as bright as ArcLight — Gong 2014, Main text
• MacQ-mOrange2 is 3× as bright as ArcLight — Gong 2014, Main text
• Marina is 0.677× as bright as ArcLight — Platisa 2017, Main text
• Mermaid is 1.286× as bright as EGFP — Tsutsui 2008, Main text
• nirButterfly is 486× as bright as Arch — Monakhov 2020, Table 1
• nirButterfly is 7× as bright as Archon1 — Song 2024, Fig. 2b
• nirButterfly is 3.17× as bright as SomArchon — Monakhov 2020, Fig. S2
• NovArch is 3× as bright as Archon1 — Chien 2021, Fig. S1C
• pAce is 0.7× as bright as Ace2N-mNeon — Kannan 2022, Fig. S11A
• pAce is 28× as bright as ASAP4e — Lee 2024, Table S4
• pAce is 0.7692307692307692× as bright as ElectraON — Zhang 2025, Fig. 1e
• pAce is 2.380952380952381× as bright as Jarvis — Grimm 2026, Fig. 1C
• pAceR is 0.7× as bright as VARNAM2 — Kannan 2022, Fig. S11A
• paQuasAr3 is 2× as bright as QuasAr2 — Adam 2019, Ext. Fig. 3f
• paQuasAr3 is 0.0017985611510791368× as bright as ASAP3 — Villette 2019, Table 1
• Positron is 0.59× as bright as Voltron — Abdelfattah 2020, Fig. 2b
• Positron is 3.5× as bright as ASAP4e — Lee 2024, Table S4
• QuasAr1 is 15.2× as bright as Arch — Hochbaum 2014, Table S2
• QuasAr1 is 20× as bright as Arch — Monakhov 2020, Table 1
• QuasAr2 is 3.4× as bright as Arch — Hochbaum 2014, Table S2
• QuasAr2 is 0.36× as bright as Archon1 — Piatkevich 2018, Fig. 2b
• QuasAr2 is 1× as bright as Archer1 — Monakhov 2020, Table 1
• QuasAr2 is 0.12484394506866417× as bright as Archon2 — Piatkevich 2018, Fig. 2b
• QuasAr2 is 0.0017985611510791368× as bright as ASAP3 — Villette 2019, Table 1
• QuasAr2 is 0.5× as bright as paQuasAr3 — Adam 2019, Ext. Fig. 3f
• QuasAr6a is 1.7× as bright as Archon1 — Tian 2023, Ext. Fig. 4b
• QuasAr6b is 2× as bright as Archon1 — Tian 2023, Ext. Fig. 4b
• rEstus is 0.775× as bright as EGFP — Rühl 2024, Fig. 1G
• rEstus is 2.07× as bright as ASAP3 — Rühl 2024, Table 1
• rEstus is 1.71× as bright as ASAP3 — Nair 2025, Table S1
• rEstus is 0.9569377990430623× as bright as rEstus-NI — Nair 2025, Fig. 2C
• rEstus is 1.669449081803005× as bright as rEstus2s — Rühl 2026, Table S1
• rEstus-NI is 1.045× as bright as rEstus — Nair 2025, Fig. 2C
• rEstus-NI is 1.788× as bright as ASAP3 — Nair 2025, Fig. 2C
• rEstus2s is 0.599× as bright as rEstus — Rühl 2026, Table S1
• rEstus2s is 1.078× as bright as ASAP3 — Rühl 2026, Table S1
• rEstus2s is 0.738× as bright as JEDI-1P — Rühl 2026, Table S1
• SomArchon is 28× as bright as Arch — Monakhov 2020, Table 1
• SomArchon is 0.31545741324921134× as bright as nirButterfly — Monakhov 2020, Fig. S2
• VADER1 is 0.85× as bright as FlicR2 — Yang 2026, Fig. 2C
• VARNAM is 2286× as bright as Arch — Monakhov 2020, Table 1
• VARNAM is 0.71× as bright as Ace2N-mNeon — Kannan 2018, Fig. 1j
• VARNAM is 0.6× as bright as Cepheid1b — Han 2023, Fig. S5
• VARNAM is 2.32× as bright as Cepheid1s — Han 2023, Fig. S5
• VARNAM is 1× as bright as VARNAM2 — Kannan 2022, Fig. S11A
• VARNAM2 is 1× as bright as VARNAM — Kannan 2022, Fig. S11A
• VARNAM2 is 1.4285714285714286× as bright as pAceR — Kannan 2022, Fig. S11A
• Vega is 2× as bright as Ace2N-mNeon2 — Cao 2025, Fig. 1B
• Vega is 8× as bright as JEDI-1P — Cao 2025, Fig. 1B
• Voltron is 4× as bright as Ace2N-mNeon — Abdelfattah 2019, Fig. 1F
• Voltron is 5.2631578947368425× as bright as HVI — Liu 2021, Table S4
• Voltron is 1.6949152542372883× as bright as Positron — Abdelfattah 2020, Fig. 2b
• Voltron is 1.5384615384615383× as bright as Voltron2 — Abdelfattah 2023, Fig. 2H
• Voltron2 is 0.65× as bright as Voltron — Abdelfattah 2023, Fig. 2H
• VSFP-Butterfly is 1.74× as bright as EGFP — doi:EC×QY: mCitrine EC=77000, QY=0.76 vs EGFP EC=56000, QY=0.60
• VSFP2.3 is 1.74× as bright as EGFP — doi:Estimated from Citrine QY=0.76, EC=77000 vs EGFP QY=0.60, EC=56000, FPbase
• VSFP3.1 is 0.79× as bright as EGFP — doi:Estimated from Cerulean EC=43000, QY=0.62 vs EGFP EC=56000, QY=0.60, FPbase

Data and citation

• llms.txt — summary for language models
• llms-full.txt — every sensor in full (Markdown)
• gevis.json — all values and measurements with sources (JSON)
• gevis.csv — one row per sensor (CSV)

Cite GEVIBench as: Adam E. Cohen and Daozheng Gong (2026). Voltage imaging: A practical guide. Neuron. https://doi.org/10.1016/j.neuron.2026.08.002

Generated from the GEVIBench database on 2026-10-12.