GEVIBench › Family tree

GEVI family tree

Lineage of every sensor in GEVIBench: each sensor is listed under the sensor it was engineered from, grouped by architecture branch.

Opsin › Opsin-FRET › Chemigenetic

• Voltron (2019)
  • Positron (2020)
    • Positron2 (2022)
  • Voltron2 (2023)
    • Solaris (2024)
• HVI (2021)
  • HVI+ (2025)
• 2Photron (2024)

Opsin › Opsin-FRET

• CAESR (2014)
• MacQ-mCitrine (2014)
• MacQ-mOrange2 (2014)
• Ace2N-mNeon (2015)
  • VARNAM (2018)
    • VARNAM2 (2022)
      • pAceR (2022)
    • Cepheid1b (2023)
      • Vega (2025)
    • Cepheid1s (2023)
  • Ace2N-mNeon2 (2022)
    • pAce (2022)
      • ElectraON (2025)
      • Jarvis (2026)
    • ElectraOFF (2025)

Opsin › Opsin-Fluorescent

• PROPS (2011)
• Arch (2012)
  • Archer1 (2014)
  • Archer2 (2014)
  • QuasAr1 (2014)
  • QuasAr2 (2014)
    • Archon1 (2018)
      • SomArchon (2019)
      • QuasAr6a (2023)
      • QuasAr6b (2023)
    • Archon2 (2018)
    • paQuasAr3 (2019)
      • NovArch (2021)

VSD › VSD-FRET

• VSFP1 (2001)
  • VSFP2.1 (2007)
    • VSFP2.3 (2008)
    • VSFP3.1 (2009)
    • VSFP-Butterfly (2012)
      • nirButterfly (2020)
    • AMBER (2021)
• Mermaid (2008)
  • Mermaid2 (2013)
• LOTUS-V (2017)

VSD › VSD-single

• ArcLight (2012)
  • Bongwoori (2015)
    • Bongwoori-R3 (2017)
  • Marina (2017)
  • HArclight1 (2021)
• ElectricPk (2012)
  • FlicR1 (2016)
    • FlicR2 (2018)
• ASAP1 (2014)
  • ASAP2s (2017)
    • ASAP3 (2019)
      • ASAP4b (2023)
        • ASAP6c (2026)
      • ASAP4e (2023)
      • ASAP5 (2024)
        • ASAP7y (2026)
      • rEstus (2024)
        • rEstus-NI (2025)
        • rEstus2s (2026)
    • JEDI-2P (2022)
      • FORCE1s (2026)
      • JEDI-3hyp (2026)
      • JEDI-3sub (2026)
      • VADER1 (2026)
    • JEDI-1P (2023)
  • HASAP1 (2021)
• Pado (2016)

Cross-branch relationships

• ArcLight is also related to Mermaid
• CAESR is also related to Arch
• HVI is also related to Ace2N-mNeon
• Positron2 is also related to pAce
• Solaris is also related to Cepheid1b
• Voltron is also related to Ace2N-mNeon

Data and citation

• llms.txt — summary for language models
• llms-full.txt — every sensor in full (Markdown)
• gevis.json — all values and measurements with sources (JSON)
• gevis.csv — one row per sensor (CSV)

Cite GEVIBench as: Adam E. Cohen and Daozheng Gong (2026). Voltage imaging: A practical guide. Neuron. https://doi.org/10.1016/j.neuron.2026.08.002

Generated from the GEVIBench database on 2026-10-12.